Loughborough University
Leicestershire, UK
LE11 3TU
+44 (0)1509 263171
Loughborough University

Loughborough University Research Publications


Publications for Jonathan Folland

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Journal Articles

Miller, R, Balshaw, TG, Massey, GJ, Maeo, S, Lanza, MB, Johnston, M, Allen, S, Folland, J (2020) The muscle morphology of elite sprint running, Medicine and Science in Sports and Exercise, ISSN: 0195-9131. DOI: 10.1249/MSS.0000000000002522.

Kordi, M, Folland, J, Goodall, S, Menzies, C, Patel, TS, Evans, M, Thomas, K, Howatson, G (2020) Cycling-specific isometric resistance training improves peak power output in elite sprint cyclists, Scandinavian Journal of Medicine and Science in Sports, 30(9), pp.1594-1604, ISSN: 0905-7188. DOI: 10.1111/sms.13742.

Maden-Wilkinson, T, Balshaw, T, Massey, G, Folland, J (2020) What makes long-term resistance-trained individuals so strong? A comparison of skeletal muscle morphology, architecture, and joint mechanics, Journal of Applied Physiology, 128(4), pp.1000-1011, ISSN: 8750-7587. DOI: 10.1152/japp.hysiol.00224.2019.

Del Vecchio, A, Negro, F, Holobar, A, Casolo, A, Folland, JP, Felici, F, Farina, D (2020) Direct translation of findings in isolated animal preparations to in vivo human motoneuron behaviour is challenging, The Journal of Physiology, 598(5), pp.1111-1112, ISSN: 0022-3751. DOI: 10.1113/JP279384.

Kordi, M, Folland, J, Goodall, S, Haralabidis, N, Maden-Wilkinson, T, Patel, TS, Leeder, J, Barratt, P, Howatson, G (2019) Mechanical and morphological determinants of peak power output in elite cyclists, Scandinavian Journal of Medicine and Science in Sports, 30(2), pp.227-237, ISSN: 0905-7188. DOI: 10.1111/sms.13570.

Kordi, M, Folland, J, Goodall, S, Barratt, P, Howatson, G (2019) Isovelocity vsIsoinertial Sprint Cycling Tests for Power- And Torque-cadence Relationships, International Journal of Sports Medicine, 40(14), pp.897-902, ISSN: 0172-4622. DOI: 10.1055/a-0989-2387.

Lanza, MB, Balshaw, T, Folland, J (2019) Is the joint-angle specificity of isometric resistance training real? And if so, does it have a neural basis?, European Journal of Applied Physiology, 119(11-12), pp.2465-2476, ISSN: 1439-6319. DOI: 10.1007/s00421-019-04229-z.

Black, MI, Allen, S, Forrester, S, Folland, J (2019) The anthropometry of economical running, Medicine and Science in Sports and Exercise, 52(3), pp.762-770, ISSN: 0195-9131. DOI: 10.1249/MSS.0000000000002158.

Hartley, C, Folland, J, Kerslake, R, Brooke-Wavell, K (2019) High-impact exercise increased femoral neck bone density with no adverse effects on imaging markers of knee osteoarthritis in postmenopausal women, Journal of Bone and Mineral Research, 35(1), pp.53-63, ISSN: 0884-0431. DOI: 10.1002/jbmr.3867.

Lanza, MB, Balshaw, T, Folland, J (2019) Explosive strength: effect of knee-joint angle on functional, neural, and intrinsic contractile properties, European Journal of Applied Physiology, 119(8), DOI: 10.1007/s00421-019-04163-0.

Kordi, M, Folland, J, Goodall, S, Barratt, PA, Howatson, G (2019) Reliabilty of traditional and task specific reference tasks to assess peak muscle activation during two different sprint cycling tests, Journal of Electromyography and Kinesiology, 46, pp.41-48, ISSN: 1050-6411. DOI: 10.1016/j.jelekin.2019.03.008.

Vecchio, AD, Negro, F, Holobar, A, Casolo, A, Folland, J, Felici, F, Farina, D (2019) You are as fast as your motor neurons: speed of recruitment and maximal discharge of motor neurons determine the maximal rate of force development in humans, Journal of Physiology, ISSN: 0022-3751. DOI: 10.1113/JP277396.

Tillin, NA, Pain, M, Folland, J (2018) Contraction speed and type influences rapid utilisation of available muscle force: neural and contractile mechanisms, Journal of Experimental Biology, 221(24), jeb193367, ISSN: 0022-0949. DOI: 10.1242/jeb.193367.

Balshaw, T, Massey, GJ, Maden-Wilkinson, TM, Lanza, MB, Folland, J (2018) Neural adaptations after 4 years vs 12 weeks of resistance training vs untrained, Scandinavian Journal of Medicine and Science in Sports, 29(3), pp.348-359, ISSN: 0905-7188. DOI: 10.1111/sms.13331.

Massey, GJ, Balshaw, T, Maden-Wilkinson, TM, Tillin, NA, Folland, J (2018) Tendinous tissue adaptation to explosive- vs. sustained-contraction strength training, Frontiers in Physiology, 9(SEP), DOI: 10.3389/fphys.2018.01170.

Massey, GJ, Balshaw, TG, Maden-Wilkinson, TM, Folland, JP (2018) Tendinous tissue properties after short and long-term functional overload: Differences between controls, 12 weeks and 4 years of resistance training., Acta Physiol (Oxf), DOI: 10.1111/apha.13019.

Malcolm, RA, Cooper, SB, Folland, J, Tyler, CJ, Sunderland, C (2018) Passive heat exposure alters perception and executive function, Frontiers in Physiology, 9(MAY), DOI: 10.3389/fphys.2018.00585.

Shaw, AJ, Ingham, SA, Folland, J (2018) The efficacy of downhill running as a method to enhance running economy in trained distance runners, European Journal of Sport Science, ISSN: 1746-1391. DOI: 10.1080/17461391.2018.1449892.

Behan, F, Maden-Wilkinson, TM, Pain, M, Folland, J (2018) Sex differences in muscle morphology of the knee flexors and knee extensors, PlosOne, 13(1), DOI: 10.1371/journal.pone.0190903.

Allison, SJ, Brooke-Wavell, K, Folland, J (2018) High and odd impact exercise training improved physical function and fall risk factors in community-dwelling older men, The Journal of Musculoskeletal and Neuronal Interactions, ISSN: 1108-7161.

Behan, F, Willis, S, Pain, M, Folland, J (2018) Effects of football simulated fatigue on neuromuscular function and whole-body response to disturbances in balance, Scandinavian Journal of Medicine and Science in Sports, ISSN: 0905-7188. DOI: 10.1111/sms.13261.

Lanza, MB, Balshaw, T, Massey, GJ, Folland, J (2018) Does normalization of voluntary EMG amplitude to MMAX account for the influence of electrode location and adiposity?, Scand J Med Sci Sports, ISSN: 0905-7188. DOI: 10.1111/sms.13270.

Behan, F, Pain, M, Folland, J (2018) Explosive voluntary torque is related to whole-body response to unexpected perturbations, Journal of Biomechanics, ISSN: 0021-9290. DOI: 10.1016/j.jbiomech.2018.09.016.

Folland, JP, Allen, SJJ, Black, MII, Handsaker, JCC, Forrester, SEE (2017) RUNNING TECHNIQUE IS AN IMPORTANT COMPONENT OF RUNNING ECONOMY AND PERFORMANCE, Medicine and Science in Sports and Exercise, 49(7), pp.1412-1423, ISSN: 1530-0315. DOI: 10.1249/MSS.0000000000001245.

Lanza, MB, Balshaw, TG, Folland, JP (2017) Do changes in neuromuscular activation contribute to the knee extensor angle-torque relationship?, Experimental Physiology, ISSN: 1469-445X. DOI: 10.1113/EP086343.

Balshaw, TG, Fry, A, Maden-Wilkinson, TM, Kong, PW, Folland, JP (2017) Reliability of quadriceps surface electromyography measurements is improved by two vs. single site recordings, European Journal of Applied Physiology, 117(6), pp.1085-1094, ISSN: 1439-6327. DOI: 10.1007/s00421-017-3595-z.

Black, MI, Handsaker, JC, Allen, SJ, Forrester, SE, Folland, JP (2017) Is There an Optimum Speed for Economical Running?, International Journal of Sports Physiology and Performance, pp.1-23, ISSN: 1555-0273. DOI: 10.1123/ijspp.2017-0015.

Massey, GJ, Balshaw, TG, Maden-Wilkinson, TM, Tillin, NA, Folland, JP (2017) The influence of patellar tendon and muscle-tendon unit stiffness on quadriceps explosive strength in man, Experimental Physiology, 102(4), pp.448-461, ISSN: 1469-445X. DOI: 10.1113/EP086190.

Balshaw, TG, Massey, GJ, Maden-Wilkinson, TM, Morales-Artacho, AJ, McKeown, A, Appleby, CL, Folland, JP (2017) Changes in agonist neural drive, hypertrophy and pre-training strength all contribute to the individual strength gains after resistance training, European Journal of Applied Physiology, 117(4), pp.631-640, ISSN: 1439-6327. DOI: 10.1007/s00421-017-3560-x.

Fry, A, Mullinger, KJ, O'Neill, GC, Brookes, MJ, Folland, JP (2016) The effect of physical fatigue on oscillatory dynamics of the sensorimotor cortex., Acta Physiologica, DOI: 10.1111/apha.12843.

Fry, A, Mullinger, KJ, O'Neill, GC, Barratt, EL, Morris, PG, Bauer, M, Folland, JP, Brookes, MJ (2016) Modulation of post-movement beta rebound by contraction force and rate of force development, Human Brain Mapping, 37(7), pp.2493-2511, ISSN: 1065-9471. DOI: 10.1002/hbm.23189.

Balshaw, TG, Massey, GJ, Maden-Wilkinson, TM, Tillin, NA, Folland, JP (2016) Training-specific functional, neural, and hypertrophic adaptations to explosive-vs. sustained-contraction strength training, Journal of Applied Physiology, 120(11), pp.1364-1373, ISSN: 8750-7587. DOI: 10.1152/japp.hysiol.00091.2016.

Maffiuletti, NA, Aagaard, P, Blazevich, AJ, Folland, J, Tillin, N, Duchateau, J (2016) Rate of force development: physiological and methodological considerations, European Journal of Applied Physiology, 116(6), pp.1091-1116, ISSN: 1439-6319. DOI: 10.1007/s00421-016-3346-6.

Evangelidis, PE, Massey, GJ, Pain, MTG, Folland, JP (2016) Strength and size relationships of the quadriceps and hamstrings with special reference to reciprocal muscle balance, European Journal of Applied Physiology, 116(3), pp.593-600, ISSN: 1439-6319. DOI: 10.1007/s00421-015-3321-7.

Evangelidis, P, Massey, GJ, Ferguson, R, Wheeler, P, Pain, M, Folland, J (2016) The functional significance of hamstrings composition: is it really a ‘fast’ muscle group?, Scandinavian Journal of Medicine and Science in Sports, ISSN: 0905-7188. DOI: 10.1111/sms.12786.

Handsaker, JC, Forrester, S, Folland, J, Black, M, Allen, S (2016) A kinematic algorithm to identify gait events during running at different speeds and with different footstrike types, Journal of Biomechanics, ISSN: 1873-2380. DOI: 10.1016/j.jbiomech.2016.10.013.

Massey, G, Evangelidis, P, Folland, J (2015) Influence of contractile force on the architecture and morphology of the quadriceps femoris, Experimental Physiology, 100(11), pp.1342-1351, ISSN: 0958-0670. DOI: 10.1113/ep085360.

Buckthorpe, M, Erskine, RM, Fletcher, G, Folland, JP (2015) Task-specific neural adaptations to isoinertial resistance training, Scandinavian Journal of Medicine & Science in Sports, 25(5), pp.640-649, ISSN: 0905-7188. DOI: 10.1111/sms.12292.

EVANGELIDIS, PE, MASSEY, GJ, PAIN, MTG, FOLLAND, JP (2015) Biceps Femoris Aponeurosis Size, Medicine & Science in Sports & Exercise, 47(7), pp.1383-1389, ISSN: 0195-9131. DOI: 10.1249/mss.0000000000000550.

Shaw, AJ, Ingham, SA, Atkinson, G, Folland, JP (2015) The Correlation between Running Economy and Maximal Oxygen Uptake: Cross-Sectional and Longitudinal Relationships in Highly Trained Distance Runners, PLOS ONE, 10(4), pp.e0123101-e0123101, DOI: 10.1371/journal.pone.0123101.

Hannah, R, Folland, JP, Smith, SL, Minshull, C (2015) Explosive hamstrings-to-quadriceps force ratio of males versus females, European Journal of Applied Physiology, 115(4), pp.837-847, ISSN: 1439-6319. DOI: 10.1007/s00421-014-3063-y.

Evangelidis, PE, Pain, MTG, Folland, J (2015) Angle-specific hamstring-to-quadriceps ratio: A comparison of football players and recreationally active males, Journal of Sports Sciences, 33(3), pp.309-319, ISSN: 0264-0414. DOI: 10.1080/02640414.2014.942680.

Hannah, R and Folland, JP (2015) Muscle-tendon unit stiffness does not independently affect voluntary explosive force production or muscle intrinsic contractile properties, Applied Physiology, Nutrition, and Metabolism, 40(1), pp.87-95, ISSN: 1715-5312. DOI: 10.1139/apnm-2014-0343.

Allison, SJ, Poole, KES, Treece, GM, Gee, AH, Tonkin, C, Rennie, WJ, Folland, J, Summers, GD, Brooke-Wavell, K (2015) The influence of high impact exercise on cortical and trabecular bone mineral content and 3D distribution across the proximal femur in older men: a randomised controlled unilateral intervention, Journal of bone and mineral research : the official journal of the American Society for Bone and Mineral Research, ISSN: 0884-0431. DOI: 10.1002/jbmr.2499.

Haider, G and Folland, JP (2014) Nitrate supplementation enhances the contractile properties of human skeletal muscle., Med Sci Sports Exerc, 46(12), pp.2234-2243, DOI: 10.1249/MSS.0000000000000351.

Folland, JP, Buckthorpe, MW, Hannah, R (2014) Human capacity for explosive force production: Neural and contractile determinants, Scandinavian Journal of Medicine & Science in Sports, 24(6), pp.894-906, ISSN: 0905-7188. DOI: 10.1111/sms.12131.

SHAW, AJ, INGHAM, SA, FOLLAND, JP (2014) The Valid Measurement of Running Economy in Runners, Medicine & Science in Sports & Exercise, 46(10), pp.1968-1973, ISSN: 0195-9131. DOI: 10.1249/mss.0000000000000311.

Fry, A, Vogt, T, Folland, JP (2014) Does sensorimotor cortex activity change with quadriceps femoris torque output? A human electroencephalography study, Neuroscience, 275, pp.540-548, ISSN: 0306-4522. DOI: 10.1016/j.neuroscience.2014.06.044.

Fry, A and Folland, JP (2014) Prolonged infrapatellar tendon vibration does not influence quadriceps maximal or explosive isometric force production in man, European Journal of Applied Physiology, 114(8), pp.1757-1766, ISSN: 1439-6319. DOI: 10.1007/s00421-014-2904-z.

Buckthorpe, M, Pain, MTG, Folland, JP (2014) Central fatigue contributes to the greater reductions in explosive than maximal strength with high-intensity fatigue., Exp Physiol, 99(7), pp.964-973, DOI: 10.1113/expphysiol.2013.075614.

Buckthorpe, M, Pain, MTG, Folland, JP (2014) Central fatigue contributes to the greater reductions in explosive than maximal strength with high-intensity fatigue, Experimental Physiology, 99(7), pp.964-973, ISSN: 0958-0670. DOI: 10.1113/expphysiol.2013.075614.

Erskine, RM, Fletcher, G, Folland, JP (2014) The contribution of muscle hypertrophy to strength changes following resistance training, European Journal of Applied Physiology, 114(6), pp.1239-1249, ISSN: 1439-6319. DOI: 10.1007/s00421-014-2855-4.

Fry, A and Folland, JP (2014) Prolonged infrapatellar tendon vibration does not influence quadriceps maximal or explosive isometric force production in man, European Journal of Applied Physiology, ISSN: 1439-6319. DOI: 10.1007/s00421-014-2904-z.

Shaw, AJ, Ingham, SA, Folland, JP (2014) The Relationship Between Running Economy and Maximal Oxygen Uptake in Trained Distance Runners, Medicine & Science in Sports & Exercise, 46, pp.725-725, ISSN: 0195-9131. DOI: 10.1249/01.mss.0000495662.42086.fd.

Shaw, AJ, Ingham, SA, Folland, JP (2014) The valid measurement of running economy in runners, Medicine and Science in Sports and Exercise, ISSN: 0195-9131. DOI: 10.1249/MSS.0000000000000311.

Tillin, NA and Folland, JP (2014) Maximal and explosive strength training elicit distinct neuromuscular adaptations, specific to the training stimulus, European Journal of Applied Physiology, 114(2), pp.365-374, ISSN: 1439-6319. DOI: 10.1007/s00421-013-2781-x.

Hannah, R, Minshull, C, Smith, SL, Folland, JP (2014) Longer electromechanical delay impairs hamstrings explosive force versus quadriceps., Med Sci Sports Exerc, 46(5), pp.963-972, DOI: 10.1249/MSS.0000000000000188.

Erskine, RM, Fletcher, G, Folland, JP (2014) The contribution of muscle hypertrophy to strength changes following resistance training, European Journal of Applied Physiology, pp.1-11, ISSN: 1439-6319. DOI: 10.1007/s00421-014-2855-4.

Haider, G and Folland, JP (2014) Nitrate supplementation enhances the contractile properties of human skeletal muscle, Medicine and Science in Sports and Exercise, 46(12), pp.2234-2243, ISSN: 0195-9131. DOI: 10.1249/MSS.0000000000000351.

Shaw, AJ, Ingham, SA, Fudge, BW, Folland, JP (2013) The reliability of running economy expressed as oxygen cost and energy cost in trained distance runners., Appl Physiol Nutr Metab, 38(12), pp.1268-1272, DOI: 10.1139/apnm-2013-0055.

Tillin, NA, Pain, MTG, Folland, JP (2013) Identification of contraction onset during explosive contractions. Response to Thompson et al. "Consistency of rapid muscle force characteristics: Influence of muscle contraction onset detection methodology" [J Electromyogr Kinesiol 2012;22(6):893-900], Journal of Electromyography and Kinesiology, 23(4), pp.991-994, ISSN: 1050-6411. DOI: 10.1016/j.jelekin.2013.04.015.

Allison, SJ, Brooke-Wavell, K, Folland, JP (2013) Multiple joint muscle function with ageing: the force-velocity and power-velocity relationships in young and older men., Aging Clin Exp Res, 25(2), pp.159-166, DOI: 10.1007/s40520-013-0024-y.

Allison, SJ, Folland, JP, Rennie, WJ, Summers, GD, Brooke-Wavell, K (2013) High impact exercise increased femoral neck bone mineral density in older men: a randomised unilateral intervention., Bone, 53(2), pp.321-328, DOI: 10.1016/j.bone.2012.12.045.

Hannah, R, Minshull, C, Folland, JP (2013) Whole-body vibration does not influence knee joint neuromuscular function or proprioception, Scandinavian Journal of Medicine and Science in Sports, 23(1), pp.96-104, ISSN: 0905-7188. DOI: 10.1111/j.1600-0838.2011.01361.x.

Tillin, NA, Pain, MTG, Folland, J (2013) Explosive force production during isometric squats correlates with athletic performance in rugby union players., J Sports Sci, 31(1), pp.66-76, DOI: 10.1080/02640414.2012.720704.

Buckthorpe, MW, Pain, MTG, Folland, JP (2013) Bilateral deficit in explosive force production is not caused by changes in agonist neural drive., PLoS One, 8(3), e57549, DOI: 10.1371/journal.pone.0057549.

Folland, JP, Buckthorpe, MW, Hannah, R (2013) Human capacity for explosive force production: Neural and contractile determinants, Scandinavian Journal of Medicine and Science in Sports, ISSN: 0905-7188. DOI: 10.1111/sms.12131.

Tillin, NA and Folland, JP (2013) Maximal and explosive strength training elicit distinct neuromuscular adaptations, specific to the training stimulus, European Journal of Applied Physiology, pp.1-10, ISSN: 1439-6319. DOI: 10.1007/s00421-013-2781-x.

Buckthorpe, MW, Hannah, R, Pain, TG, Folland, JP (2012) Reliability of neuromuscular measurements during explosive isometric contractions, with special reference to electromyography normalization techniques., Muscle Nerve, 46(4), pp.566-576, DOI: 10.1002/mus.23322.

Erskine, RM, Fletcher, G, Hanson, B, Folland, JP (2012) Whey protein does not enhance the adaptations to elbow flexor resistance training., Med Sci Sports Exerc, 44(9), pp.1791-1800, DOI: 10.1249/MSS.0b013e318256c48d.

Tillin, NA, Pain, MTG, Folland, JP (2012) Contraction type influences the human ability to use the available torque capacity of skeletal muscle during explosive efforts., Proc Biol Sci, 279(1736), pp.2106-2115, DOI: 10.1098/rspb.2011.2109.

Folland, JP, Mc Cauley, TM, Phypers, C, Hanson, B, Mastana, SS (2012) The relationship of testosterone and AR CAG repeat genotype with knee extensor muscle function of young and older men., Exp Gerontol, 47(6), pp.437-443, DOI: 10.1016/j.exger.2012.03.013.

Tillin, NA, Pain, MTG, Folland, JP (2012) Short-term training for explosive strength causes neural and mechanical adaptations., Exp Physiol, 97(5), pp.630-641, DOI: 10.1113/expphysiol.2011.063040.

Folland, JP, Mc Cauley, TM, Phypers, C, Hanson, B, Mastana, SS (2012) Relationship of 2D:4D finger ratio with muscle strength, testosterone, and androgen receptor CAG repeat genotype, American Journal of Physical Anthropology, 148(1), pp.81-87, ISSN: 0002-9483. DOI: 10.1002/ajpa.22044.

Folland, JP and Fry, A (2012) Neual Drive during Explosive Force Production exceeds that at Maximum Force, MEDICINE AND SCIENCE IN SPORTS AND EXERCISE, 44, pp.547-547, ISSN: 0195-9131.

Buckthorpe, M, Morris, J, Folland, JP (2012) Validity of vertical jump measurement devices, Journal of Sports Sciences, 30(1), pp.63-69, ISSN: 0264-0414. DOI: 10.1080/02640414.2011.624539.

Hannah, R, Minshull, C, Buckthorpe, MW, Folland, JP (2012) Explosive neuromuscular performance of males versus females, Experimental Physiology, 97(5), pp.618-629, ISSN: 0958-0670. DOI: 10.1113/expphysiol.2011.063420.

Folland, JP, Buckthorpe, MW, Hannah, R (2011) Neural and Contractile Determinants of Explosive Force Production, Medicine & Science in Sports & Exercise, 43(Suppl 1), pp.101-101, ISSN: 0195-9131. DOI: 10.1249/01.mss.0000402975.04207.b5.

Tillin, NA, Pain, MTG, Folland, JP (2011) Short-term unilateral resistance training affects the agonist-antagonist but not the force-agonist activation relationship, Muscle and Nerve, 43(3), pp.375-384, ISSN: 1097-4598. DOI: 10.1002/mus.21885.

Hannah, R, Minshull, C, Folland, JP (2011) Whole-body vibration does not influence knee joint neuromuscular function or proprioception, Scandinavian Journal of Medicine and Science in Sports, ISSN: 0905-7188.

Folland, JP, Haas, B, Castle, PC (2011) Strength and activation of the knee musculature in Parkinson's disease: Effect of medication, NEUROREHABILITATION, 29(4), pp.405-411, ISSN: 1053-8135. DOI: 10.3233/NRE-2011-0719.

Neville, V, Gant, N, Folland, JP (2010) Thermoregulatory demands of Elite Professional America's Cup Yacht Racing, Scandinavian Journal of Medicine & Science in Sports, 20(3), pp.475-484, DOI: 10.1111/j.1600-0838.2009.00952.x.

McCauley, TM, Mastana, S, Folland, JP (2010) ACE I/D and ACTN3 R/X polymorphisms and muscle function and muscularity of older Caucasian men, European Journal of Applied Physiology, 109(2), pp.269-277, ISSN: 1439-6319. DOI: 10.1007/s00421-009-1340-y.

Tillin, NA, Jimenez Reyes, P, Pain, MTG, Folland, JP (2010) Neuromuscular Performance of Explosive Power Athletes versus Untrained Individuals, Medicine & Science in Sports & Exercise, 42(4), pp.781-790, ISSN: 0195-9131. DOI: 10.1249/MSS.0b013e3181be9c7e.

Neville, V, Pain, MTG, Kantor, J, Folland, JP (2010) Influence of Crank Length and Crank-Axle Height on Standing Arm-Crank (Grinding) Power, Medicine & Science in Sports & Exercise, 42(2), pp.381-387, ISSN: 0195-9131. DOI: 10.1249/MSS.0b013e3181b46f3a.

Neville, V, Zaher, N, Pain, MTG, Folland, JP (2009) Lower Limb Influence on Standing Arm-cranking (`grinding'), International Journal of Sports Medicine, 30(10), pp.713-718, ISSN: 0172-4622. DOI: 10.1055/s-0029-1231044.

Neville, V, Calefato, J, Perez Encinas, C, Rodilla Sala, E, Rada Ruiz, S, Dorochenko, P, Folland, JP (2009) America's Cup yacht racing: Race analysis and physical characteristics of the athletes, Journal of Sports Sciences,, 27(9), pp.915-923, ISSN: 0264-0414. DOI: 10.1080/02640410902946485.

Neville, V, Pain, MTG, Folland, JP (2009) Aerobic power and peak power of elite America's Cup sailors., Eur J Appl Physiol, 106(1), pp.149-157, DOI: 10.1007/s00421-009-1002-0.

Neville, V and Folland, JP (2009) The epidemiology and aetiology of injuries in sailing, Sports Medicine, 39(2), pp.129-145, DOI: 10.2165/00007256-200939020-00003.

McCauley, TM, Mastana, S, Hossack, J, Macdonald, M, Folland, JP (2009) Human angiotensin-converting enzyme I/D and alpha-actinin 3 R577X genotypes and muscle functional and contractile properties, Experimental Physiology, 94(1), pp.81-89, ISSN: 0958-0670. DOI: 10.1113/expphysiol.2008.043075.

Folland, JP (2009) Measurement of maximum muscle activation with the interpolated twitch technique, Journal of Applied Physiology, 107, pp.365-366.

Williams, AG and Folland, JP (2008) Reply from Alun G. Williams and Jonathan P. Folland, Journal of Physiology, 586(12), pp.3019-3020, ISSN: 0022-3751. DOI: 10.1113/jphysiol.2008.155895.

Folland, JP, Mc Cauley, TM, Williams, AG (2008) Allometric scaling of strength measurements to body size., Eur J Appl Physiol, 102(6), pp.739-745, ISSN: 1439-6319. DOI: 10.1007/s00421-007-0654-x.

Folland, J and Morris, B (2008) Variable-cam resistance training machines: do they match the angle - torque relationship in humans?, J Sports Sci, 26(2), pp.163-169, ISSN: 0264-0414. DOI: 10.1080/02640410701370663.

Williams, AG and Folland, JP (2008) Similarity of polygenic profiles limits the potential for elite human physical performance, Journal of Physiology, 586(1), pp.113-121, ISSN: 1469-7793. DOI: 10.1113/jphysiol.2007.141887.

Williams, AG and Folland, JP (2008) The genetic basis of human athletic performance. Why are psychological components so often overlooked? Reply from the authors, Journal of Physiology, 586, pp.3019-3020.

Allison, SJ, Bailey, DM, Folland, JP (2008) Prolonged static stretching does not influence running economy despite changes in neuromuscular function, Journal of Sports Science, 26(14), pp.1489-1495, DOI: 10.1080/02640410802392715.

Folland, JP, Wakamatsu, T, Fimland, MS (2008) The influence of maximal isometric activity on twitch and H-reflex potentiation, and quadriceps femoris performance, European Journal of Applied Physiology, 104(4), pp.739-748, DOI: 10.1007/s00421-008-0823-6.

Neville, V, Gleeson, M, Folland, JP (2008) Salivary IgA as a risk factor for upper respiratory infections in elite professional athletes, Medicine and Science in Sports and Exercise, 40(7), pp.1228-1236.

Lindh, AM, Peyrebrune, MC, Ingham, SA, Bailey, DM, Folland, JP (2007) Sodium bicarbonate supplementation improves 200m freestyle performance in elite male swimmers, International Journal of Sports Medicine, 29(6), pp.519-523, DOI: 10.1055/s-2007-989228.

Folland, JP, Stern, R, Brickley, G (2007) Sodium Phospate loading improves laboratory cycling time trial performance in trained cyclists, Journal of Science and Medicine in Sport, 11, pp.464-468, DOI: 10.1016/j.jsams.2007.04.004.

Folland, JP and Williams, AG (2007) Methodological issues with the interpolated twitch technique, Journal of Electromyography and Kinesiology, 17(3), pp.317-327, ISSN: 1050-6411. DOI: 10.1016/j.jelekin.2006.04.008.

Folland, JP and Williams, AG (2007) Calculation of muscle activation using neuromuscular electrical stimulation, Medicine and Science in Sports and Exercise, 39, p.745.

Folland, JP and Williams, AG (2007) The adaptations to strength training : morphological and neurological contributions to increased strength., Sports Med, 37(2), pp.145-168, ISSN: 0112-1642. DOI: 10.2165/00007256-200737020-00004.

Folland, JP, Rowlands, DS, Thorp, R, Walmsley, A (2006) Leg heating and cooling influences running stride parameters but not running economy, International Journal of Sports Medicine, 27, pp.771-779.

Williams, AG, Day, SH, Folland, JP, Gohlke, P, Dhamrait, S, Montgomery, HE (2005) Circulating angiotensin converting enzyme activity is correlated with muscle strength, Medicine and Science in Sports and Exercise, 37(6), pp.944-948, ISSN: 0195-9131. DOI: 10.1249/01.mss.0000166577.42935.4e.

Folland, JP, Hawker, K, Leach, B, Little, T, Jones, DA (2005) Strength training: Isometric training at a range of joint angles versus dynamic training, Journal of Sports Sciences, 23, pp.817-824.

Day, SH, Williams, C, Folland, JP, Gohlke, P, Williams, AG (2004) The acute effects of exercise and glucose ingestion on circulating angiotensin-converting enzyme in humans, European Journal of Applied Physiology, 92(4-5), pp.579-583, ISSN: 1439-6327. DOI: 10.1007/s00421-004-1078-5.

Folland, JP, Irish, CS, Roberts, JC, Tarr, JE, Jones, DA (2002) Fatigue is not a necessary stimulus for strength gains during resistance training, British Journal of Sports Medicine, 36(5), pp.370-373, ISSN: 1473-0480. DOI: 10.1136/bjsm.36.5.370.

Folland, JP, Chong, J, Copeman, EM, Jones, DA (2001) Acute muscle damage as a stimulus for training-induced gains in strength., Med Sci Sports Exerc, 33(7), pp.1200-1205, ISSN: 0195-9131. DOI: 10.1097/00005768-200107000-00019.

Folland, JP, Maas, H, Jones, DA (2000) The influence of nitric oxide on in-vivo human skeletal muscle properties, Acta Physiologica Scandanavia, 169, pp.141-148, ISSN: 0001-6772.

Folland, JP, Leach, B, Little, T, Hawker, K, Myerson, S, Montgomery, H, Jones, DA (2000) Angiotensin converting enzyme genotype affects the response of skeletal muscle to functional overload, Experimental Physiology, 85(5), pp.575-579.

Folland, JP, Hawker, K, Leach, B, Little, T, Myerson, S, Jones, DA (2000) ACE genotype affects the strength training response, Journal of Sports Sciences, 18(7), p.536, ISSN: 0264-0414.

Harrison, AJ and Folland, JP (1998) The validity and reliability of optical dynamic pedobarograph data, British Journal of Podiatry, 53(1), pp.6-10.

Harrison, AJ and Folland, JP (1997) Investigation of gait protocols for plantar pressure measurement of non-pathological subjects using a dynamic pedobarograph, Gait & Posture, 6, pp.50-55.



Conferences

Hartley, C, Folland, J, Brooke-Wavell, K, Kerslake, R (2019) High impact exercise increased femoral neck bone density with no adverse effects on imaging markers of knee osteoarthritis in postmenopausal women [Abstract]. In 2019 Annual Meeting of the American Society for Bone and Mineral Research; Journal of Bone and Mineral Research, Orlando, Florida, USA, pp.107-107, DOI: 10.1002/jbmr.3936.

Hartley, C, Folland, JP, Brooke-Wavell, K, Kerslake, R (2019) High impact exercise increased femoral neck bone density with no adverse effects on imaging markers of knee osteoarthritis in postmenopausal women. In , JOURNAL OF BONE AND MINERAL RESEARCH, pp.107-107.

Howatson, G, Kordi, M, Goodall, S, Haralabidis, N, Patel, T, Maden-Wilkinson, T, Folland, J (2019) Physiological Determinants of Peak Power Output in Elite Cyclists. In , MEDICINE AND SCIENCE IN SPORTS AND EXERCISE, pp.638-638, DOI: 10.1249/01.mss.0000562406.76129.38.

Hartley, C, Kerslake, RW, Folland, J, Brooke-Wavell, K (2018) The reproducibility of T2 relaxation time measurement of knee cartilage in women aged over 50 [Abstract]. In , Liverpool. DOI: 10.1016/j.joca.2018.02.879.

Lanza, MB, Balshaw, TB, Folland, JP (2017) MMAX Normalisation of Voluntary EMG Removes the Confounding Influences of Electrode Location and Body Fat.. In , Medicine & Science in Sports & Exercise, pp.779-779, DOI: 10.1249/01.mss.0000519076.74034.cb.

Allison, SJ, Folland, J, Brooke-Wavell, K (2016) Multidirectional hopping exercise improved balance and ankle plantarflexion strength in community-dwelling older men [Abstract]. In , Birmingham, UK, pp.666-666, DOI: 10.1007/s00198-016-3743-z.

Brooke-Wavell, K, Treece, GM, Allison, SJ, Folland, J, Rennie, WJ, Summers, GD, Gee, AH, Poole, KES (2016) Brief high impact exercise increased cortical mass and trabecular density at regions predictive of femoral neck and trochanteric fracture [Abstract]. In , Birmingham, UK, pp.626-627, DOI: 10.1007/s00198-016-3743-z.

Brooke-Wavell, K, Yang, L, Blackwell, D, Qasim, M, Allison, SJ, Rennie, WJ, Folland, J, Summers, GD, Eastell, R (2016) Increases in proximal femur bone strength estimated using finite element models from computed tomography scans following brief, high impact exercise in older men [Abstract]. In , Malaga, SPAIN. DOI: 10.1007/s00198-016-3530-x.

Behan, F, Maden-Wilkinson, TM, Pain, MTG, Folland, JP (2016) Sex Differences In Hamstrings To Quadriceps Muscle Volume Ratio. In , Medicine & Science in Sports & Exercise, pp.388-388, DOI: 10.1249/01.mss.0000486172.10588.a6.

Balshaw, TG, Massey, GJ, Maden-Wilkinson, TM, Tillin, NA, Folland, JP (2016) Twelve Weeks Of Explosive Strength Training Increases Both Maximal And Explosive Voluntary Torque Production. In , Medicine & Science in Sports & Exercise, pp.452-452, DOI: 10.1249/01.mss.0000486358.39688.0e.

Sykora, M, Chung, P, Folland, J, Halkon, B, Edirisinghe, E (2015) Advances in sports informatics research. In , Inst Teknologi Brunei, Bandar Seri Begawan, BRUNEI, pp.265-274, ISBN: 9783319131528. DOI: 10.1007/978-3-319-13153-5_26.

Folland, J and Archer, G (2014) Shoulder injuries in highly trained competitive swimmers.. In , Br J Sports Med, England, p.593, DOI: 10.1136/bjsports-2014-093494.90.

Evangelidis, P, Massey, G, Pain, M, Folland, J (2014) Is biceps femoris long head aponeurosis size a risk factor for hamstring strain injury?. In , Br J Sports Med, England, pp.591-592, DOI: 10.1136/bjsports-2014-093494.86.

Allison, S, Poole, K, Treece, G, Gee, A, Tonkin, C, Rennie, W, Folland, J, Summers, G, Brooke-Wavell, K (2014) The Influence of Exercise on the 3D distribution of Cortical and Trabecular Bone across the Proximal Femur: The HipHop Study.. In , JOURNAL OF BONE AND MINERAL RESEARCH, pp.S6-S6.

Hannah, R, Minshull, C, Smith, SL, Folland, JP (2014) Longer electromechanical delay impairs hamstrings explosive force versus quadriceps. In , Medicine and Science in Sports and Exercise, pp.963-972, DOI: 10.1249/MSS.0000000000000188.

Shaw, AJ, Ingham, SA, Fudge, BW, Folland, JP (2013) The Reliability of Running Economy Expressed as Oxygen Cost and Energy Cost in Distance Runners. In , MEDICINE AND SCIENCE IN SPORTS AND EXERCISE, pp.162-162.

Folland, JP, Fry, A, Ferguson, RA (2013) The Influence Of Skeletal Muscle MHC Composition On Explosive Force Production In Man. In , MEDICINE AND SCIENCE IN SPORTS AND EXERCISE, pp.184-184.

Allison, SJ, Folland, JP, Rennie, WJ, Summers, GD, Brooke-Wavell, K (2012) BRIEF HIGH IMPACT EXERCISE IMPROVED FEMORAL NECK BONE MINERAL DENSITY AND HIP STRUCTURAL PARAMETERS IN OLDER MEN: A RANDOMISED UNILATERAL INTERVENTION. In Osteoporosis and Bone Conference, OSTEOPOROSIS INTERNATIONAL, Manchester, ENGLAND, pp.S523-S524.

Tillin, NA, Pain, MTG, Folland, JP (2012) Training for Maximal vs. Explosive Strength Elicits Distinct Neuromuscular Adaptations. In , MEDICINE AND SCIENCE IN SPORTS AND EXERCISE, pp.87-88.

Allison, SJ, Folland, JP, Rennie, WJ, Summers, GD, Brooke-Wavell, K (2012) High Impact Unilateral Exercise Increased Femoral Neck Bone Mineral Density in Older Men. In , MEDICINE AND SCIENCE IN SPORTS AND EXERCISE, pp.707-707.

Buckthorpe, MW, Pain, MTG, Folland, JF (2012) Fatigue induces a greater decline in explosive than maximal force production with evidence for neural and contractile mechanisms. In Proceedings of the 16th Annual Congress of the European College of Sport Science, Bruges, -.

Buckthorpe, M, Pain, MTG, Folland, JF (2012) A bilateral deficit in explosive, but not maximal force production. In The Biomedical Basis of Elite Performance, Queen Elizabeth the 2nd Conference Centre, London, pp.0-0.

Buckthorpe, M, Hannah, R, Pain, MTG, Folland, JF (2011) Neural adaptations explain the greater increase in 1RM than isometric strength following short-term resistance training. In British Association of Sport & Exercise Sciences, Essex, pp.0-0.

Tillin, NA, Pain, MTG, Oguz, HE, Lewis, GA, Folland, JP (2010) A unique method of measuring dynamic rate of torque development: comparison of explosive power athletes and controls. In Proceedings of the 33rd Annual Meeting of the American Society of Biomechanics, Rhode Island, USA, pp.915-923.

Oliveira, MR, Folland, JP, Tauler, P, Allen, S, Fowkes, R, Gleeson, M (2010) A biphasic pattern of muscle enzyme efflux after eccentric exercise is associated with greater loss of muscle force and power. In Proceedings of the 15th Annual Congress of the European College of Sport Science, Antalya.

Oliveira, MR, Folland, JP, Tauler, P, Allen, S, Fowkes, R, Gleeson, M (2010) A biphasic pattern of muscle enzyme efflux after eccentric exercise is associated with greater loss of muscle force and power. In .

Tillin, NA, Pain, MTG, Curbishley, MA, Folland, JP (2010) Mechanical and neural adaptations to short-term explosive strength training. In Proceedings of the Biomechanics Interest Group of the British Association of Sport and Exercise Sciences, University of Bath, Bath, pp.915-923.

Hannah, R, Buckthorpe, M, Minshull, C, Folland, JP (2010) Explosive neuromuscular performance of the knee extensors in untrained males and females. In British Association of Sport & Exercise Sciences, ? pp.365-366.

Tillin, NA, Pain, MTG, Curbishley, MA, Folland, JP (2010) Training explosive muscle performance affects agonist activation and the mechanical properties of the muscle-tendon unit. In Proceedings of the International Society of Electromyography and Kinesiology, Aalborg, Denmark, pp.365-366.

Hannah, R, Minshull, C, Compton, G, Gleeson, N, Folland, JP (2010) The effects of a single session of whole body vibration exercise on knee joint neuromuscular function and position sense. In Proceedings of the International Society of Electromyography and Kinesiology, Aalborg, Denmark, pp.365-366.

Neville, V, Pain, MTG, Folland, JP (2009) Optimising Arm-crank Configurations: Testing the Most Powerful Upper-body Athletes in Sport. In Proceedings of the British Association of Sports and Exercise Sciences Annual Conference, Leeds, UK, pp.149-157.

Neville, V, Pain, MTG, Folland, JP (2009) Elite America's cup sailors are among the most powerful upper body trained athletes. In American College of Sports Medicine's 56th Annual Meeting, Medicine & Science in Sports and Exercise, Seattle, Washington, pp.149-157.

Tillin, NA, Pain, MTG, Folland, JP (2009) Strength training affects the agonist-antagonist but not the force-agonist activation relationship. In Proceedings of the Physiological Society 14 PC25, Journal of Physiology, Kings College, London, pp.149-157.

Tillin, N, Jimenez, P, Pain, MTG, Folland, J (2008) Differences in rate of force development and neural activation between elite explosive power athletes and controls. In Proceedings of the 13th Annual Congress of the European College of Sports Science, Estoril, Portugal, pp.575-579.

Tillin, N, Jimenez, P, Pain, MTG, Folland, J (2008) A comparison of rate of force development and neural activation between elite power athletes and controls. In Proceedings of the Biomechanics Interest Group of the British Association of Sport and Exercise Sciences, University of Wales Institute, Cardiff, p.12.

McCauley, T, Mastana, S, Folland, JP (2008) The influence of ACE I/D polymorphism on human muscle function, contractile properties and muscularity of older men. In Proceedings of the 12th Annual Congress of the European College of Sports Science, Estoril, p.153.

Neville, V, Gant, N, Folland, JP (2008) Thermoregulatory demands of elite professional America's Cup sailing. In Proceedings of the 12th Annual Congress of the European College of Sports Science, Estoril, p.497.

Neville, V, Gleeson, M, Folland, JP (2008) Salivary IgA can help to predict upper respiratory infection in athletes. In American College of Sports Medicine, Medicine and Science in Sports and Exercise, Indianapolis, S15.

McCauley, T, Mastana, S, Hossack, J, MacDonald, M, Gilmour, A, Folland, JP (2007) The influence of alpha actinin 3R/X genotype upon human muscle function and contractile properties. In Proceedings of Life Sciences, Glasgow, UK, PC432.

Folland, JP, Wakamatsu, T, Fimland, MS (2007) Post-activation potentiation: neurophysiology and strength performance. In American College of Sports Medicine, Medicine and Science in Sports and Exercise, New Orleans, pp.S303-304.

Peyrebrune, MC, Lindh, AM, Ingham, SA, Bailey, DM, Folland, JP (2007) Sodium bicarbonate supplementation improves 200m freestyle performance in elite male swimmers. In American College of Sports Medicine, Medicine and Science in Sports and Exercise, New Orleans, S218.

Williams, AG and Folland, JP (2007) An algorithm for genetic potential to express muscle strength. In Proceedings of the 12th Annual Congress of the European College of Sports Science, Jyvaskyla, Finland, pp.356-7.

Williams, AG and Folland, JP (2007) Similarity of polygenic profiles limits the potential for elite human physical performance. In Proceedings of the 12th Annual Congress of the European College of Sports Science, Jyvaskyla, Finland, pp.83-84.

Neville, V, Gleeson, M, Folland, JP (2007) Salivary IgA is a risk factor for upper respiratory infection: a longitudinal prospective study in athletes. In ACSM, Medicine and Science in Sports and Exercise, venue, pp.739-748.

McCauley, T, Mastana, S, Hossack, J, MacDonald, M, Gilmour, A, Folland, JP (2006) The influence of Angiotensin Converting Enzyme (ACE) genotype upon muscle strength and contractile properties. In British Association of Sport and Exercise Sciences, Journal of Sports Sciences, Wolverhampton, pp.270-271.

Morrisson, LLH, Peyrebrune, M, Folland, JP (2005) Resisted-swimming training improves 100m freestyle performance in elite swimmers. In British Association of Sport and Exercise Sciences, Journal of Sports Sciences, Loughborough, UK, p.1295.

Folland, JP, Rowlands, DS, Thorp, R, Walmsley, A (2005) Running economy is not affected by leg heating and cooling despite changes in running stride parameters. In Proceedings of the 10th Annual Congress of the European College of Sports Science, Belgrade, Serbia, p.161.

Folland, JP (2002) Methodological considerations with the twitch interpolation technique. In Proceedings of the 7th Annual Congress of the European College of Sports Science, Athens, Greece, p.705.

Folland, JP (2001) Sodium Phospate loading can improve laboratory 10 mile cycling performance in training cyclists. In First International Scientific Congress on Nutrition and Athletic Performance, Canadian Journal of Applied Physiology, Edmonton, Canada, S253.

Folland, JP (2000) Post contraction potentiation and the twitch interpolation technique. In Proceedings of the 5th Annual Congress of the European College of Sports Science, Jyvaskyla, Finland, p.266.

Folland, JP, Maas, H, Jones, DA (1999) The influence of nitric oxide on in-vivo human skeletal muscle properties. In American College of Sports Medicine, Medicine and Science in Sports and Exercise, Seattle, S184.

Folland, JP, Leach, B, Little, T, Hawker, K, Myerson, S, Montgomery, H, Jones, DA (1999) ACE genotype affects the strength training response. In European College of Sports Science, Journal of Sports Science, Rome, p.536.

Folland, JP, Chong, J, Copeman, EM, Jones, DA (1998) A bout of damaging eccentric work does not assist in strength training. In Proceedings of the 3rd Annual Congress of the European College of Sports Science, Manchester, p.285.

Folland, JP and Jones, DA (1998) Long-term creatine supplementation does not enhance the muscular strength of habitual trainers. In British Association of Sport and Exercise Sciences, Journal of Sports Sciences, Worcester, pp.33-34.

Folland, JP, Irish, CS, Roberts, JC, Tarr, JE, Jones, DA (1997) Muscle fatigue is not essential in strength training. In The Physiological Society, Journal of Physiology, Cambridge, 102P.

Folland, JP and Harrison, AJ (1996) The reliability of different gait protocols for foot pressure measurement. In Proceedings of In-vivo pressure measurement, Manchester, pp.83-84.



Chapters

Wilson, IS, Folland, J, McDermott, H, Munir, F (2016) White-water paddlesports medicine: Canoeing, kayaking and rafting. In Extreme Sport Medicine, © Springer, pp.289-304, ISBN: 9783319282633.

Folland, JP and Strachan, K (2013) Whitewater canoeing and rafting. In Adventure and Extreme Sports Injuries: Epidemiology, Treatment, Rehabilitation and Prevention, pp.113-141, ISBN: 9781447143628. DOI: 10.1007/978-1-4471-4363-5_6.

Folland, JP, Bailey, C, Brooke Wavell, K (2007) The importance of strength training. In Lanham-New, O'Neill, S, T, Morris, R, Skelton, D, Sutcliffe, Eds, A (ed) Managing Osteoporosis, Clinical Publishing, pp.169-184.

Jones, DA and Folland, JP (2001) Strength training in young adults. In Eds, MEA (ed) Sports Medicine for Specific Ages and Abilities, Churchill Livingstone, pp.57-64.



Other

Balshaw, T, Massey, GJ, Maden-Wilkinson, TM, Folland, J (2017) Muscle size and strength: debunking the “completely separate phenomena” suggestion, Muscle size and strength: debunking the “completely separate phenomena” suggestion. DOI: 10.1007/s00421-017-3616-y.



Getting in touch

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