Journal Articles

Guareschi, S, Mathers, K, South, J, Navarro, LM, Renals, T, Hiley, A, Antonsich, M, Bolpagni, R, Bortolus, A, Genovesi, P, Jere, A, Madzivanzira, TC, Phaka, FM, Novoa, A, Olden, JD, Saccó, M, Shackleton, RT, Vilà, M, Wood, P (2024) Framing challenges and polarized issues in invasion science: toward an interdisciplinary agenda, BioScience, ISSN: 0006-3568. DOI: 10.1093/biosci/biae084.

White, J, Seddon, E, Hill, M, Mathers, K, Bridger, M, Hannah, D, Wood, P (2024) Going to the archives: Combining palaeoecological andcontemporary data to support river restoration appraisals, River Research and Applications: an international journal devoted to river research and management, ISSN: 0886-9375.

Mathers, K, Armitage, PD, Hill, M, Bickerton, M, Mckenzie, M, Pardo, I, Tickner, D, Wood, PJ (2024) Context specific effects of substrate composition on the taxonomic and functional diversity of macroinvertebrate communities in temperate lowland streams, Ecology and Evolution, 14(8), ISSN: 2045-7758. DOI: 10.1002/ece3.70034.

Sanders, CH, Buckley, PL, Hunt, CD, Mathers, K, Mills, D (2024) Ponto-Caspian amphipod co-location with zebra mussel beds (Dreissena polymorpha) is influenced by substrate size and population source, Hydrobiologia, 851(14), ISSN: 0018-8158. DOI: 10.1007/s10750-024-05515-4.

Snåre, H, García-Girón, J, Alahuhta, J, Bini, LM, Boda, P, Bonada, N, Brasil, LS, Callisto, M, Castro, DMP, Chen, K, Csabai, Z, Datry, T, Domisch, S, Márquez, JRG, Floury, M, Friberg, N, Gill, BA, González-Trujillo, JD, Göthe, E, Haase, P, Hamada, N, Hill, MJ, Hjort, J, Juen, L, Jupke, JF, de Faria, APJ, Li, Z, Ligeiro, R, Linares, MS, Luiza-Andrade, A, Macedo, DR, Mathers, K, Mellado-Díaz, A, Milosevic, D, Moya, N, Poff, NL, Rolls, RJ, Roque, FO, Saito, VS, Sandin, L, Schäfer, RB, Scotti, A, Siqueira, T, Martins, RT, Valente-Neto, F, Wang, B, Wang, J, Xie, Z, Heino, J (2024) The relationships between biotic uniqueness and abiotic uniqueness are context dependent across drainage basins worldwide, Landscape Ecology, 39, 86, ISSN: 0921-2973. DOI: 10.1007/s10980-024-01883-3.

Mckenzie, M, Scott, J, Wood, P, Mathers, K (2024) Phenotypic plasticity of a Baetid mayfly larvae (Baetis rhodani) at sites with high levels of deposited fine sediment, Ecological Entomology, ISSN: 0307-6946. DOI: 10.1111/een.13321.

Mckenzie, M, Brooks, A, Callisto, M, Collins, AL, Durkota, JM, Death, RG, Jones, JI, Linares, MS, Matthaei, CD, Monk, WA, Murphy, JF, Wagenhoff, A, Wilkes, M, Wood, PJ, Mathers, K (2024) Freshwater invertebrate responses to fine sediment stress: A multi‐continent perspective, Global Change Biology, 30(1), ISSN: 1354-1013. DOI: 10.1111/gcb.17084.

Davis, NG, Mathers, K, Hodson, R, Matthaei, CD (2023) Monthly sampling reveals seasonal fine sediment fluctuations and riverine invertebrate community responses, Science of The Total Environment, 911(2024), 168750, ISSN: 0048-9697. DOI: 10.1016/j.scitotenv.2023.168750.

Mathers, K, Armitage, PD, Hill, M, Mckenzie, M, Pardo, I, Wood, P (2023) Seasonal variability of lotic macroinvertebrate communities at the habitat scale demonstrates the value of discriminating fine sediment fractions in ecological assessments, Ecology and Evolution, 13(10), ISSN: 2045-7758. DOI: 10.1002/ece3.10564.

Sanders, CH, Rice, S, Wood, P, Mathers, K (2023) The long-term dynamics of invasive signal crayfish forcing of fluvial sediment supply via riverbank burrowing, Geomorphology, 442, 108924, ISSN: 0169-555X. DOI: 10.1016/j.geomorph.2023.108924.

Mathers, K, Clinton, K, Constable, D, Gerrard, C, Patel, C, Wood, P (2023) Invasion dynamics of Ponto‐Caspian amphipods leads to changes in invertebrate community structure and function, Ecosphere, 14(7), e4593, ISSN: 2150-8925. DOI: 10.1002/ecs2.4593.

Mathers, K, Guareschi, S, Pattison, Z (2022) Biological invasions in rivers and associated ecosystems: New insights, challenges, and methodological advancements, River Research and Applications, 38(8), pp.1351-1355, ISSN: 1535-1459. DOI: 10.1002/rra.4038.

Kowarik, C, Martin-Creuzburg, D, Mathers, K, Weber, C, Robinson, CT (2022) Stream degradation affects aquatic resource subsidies to riparian ground-dwelling spiders, Science of The Total Environment, 855, 158658, ISSN: 0048-9697. DOI: 10.1016/j.scitotenv.2022.158658.

Mathers, K, Rice, S, Chadd, R, Wood, P (2022) Influence of invasive crayfish on fine sediment transport, ingress and bed storage in lowland rivers, River Research and Applications, 38(8), pp.1481-1494, ISSN: 1535-1459. DOI: 10.1002/rra.3995.

Mathers, K, Doretto, A, Fenoglio, S, Hill, MJ, Wood, P (2022) Temporal effects of fine sediment deposition on benthic macroinvertebrate community structure, function and biodiversity likely reflects landscape setting, Science of The Total Environment, 829, 154612, ISSN: 0048-9697. DOI: 10.1016/j.scitotenv.2022.154612.

Mathers, K, Robinson, CT, Weber, C (2022) Patchiness in flow refugia use by macroinvertebrates following an artificial flood pulse, River Research and Applications, 38(4), pp.696-707, ISSN: 1535-1459. DOI: 10.1002/rra.3941.

Hill, MJ, Greaves, HM, Sayer, CD, Hassall, C, Milin, M, Milner, VS, Marazzi, L, Hall, R, Harper, LR, Thornhill, I, Walton, R, Biggs, J, Ewald, N, Law, A, Willby, N, White, JC, Briers, RA, Mathers, K, Jeffries, MJ, Wood, P (2021) Pond ecology and conservation: research priorities and knowledge gaps, Ecosphere, 12(12), e03853, ISSN: 2150-8925. DOI: 10.1002/ecs2.3853.

Mathers, K, Guareschi, S, Patel, C, Wood, P (2021) Response of freshwater snails to invasive crayfish varies with physiochemical exposure cues and predator experience, Freshwater Biology, 67(3), pp.473-486, ISSN: 0046-5070. DOI: 10.1111/fwb.13855.

Mathers, K, Robinson, CT, Weber, C (2021) Artificial flood reduces fine sediment clogging enhancing hyporheic zone physicochemistry and accessibility for macroinvertebrates, Ecological Solutions and Evidence, 2(4), e12103, ISSN: 2688-8319. DOI: 10.1002/2688-8319.12103.

Hill, MJ, Wood, P, Mathers, K (2021) Taxonomic and functional macroinvertebrate diversity of high‐altitude ponds in the Macun Cirque, Switzerland, Aquatic Conservation: Marine and Freshwater Ecosystems, 31(11), pp.3201-3214, ISSN: 1052-7613. DOI: 10.1002/aqc.3691.

Mathers, K, Kowarik, C, Rachelly, C, Robinson, CT, Weber, C (2021) The effects of sediment traps on instream habitat and macroinvertebrates of mountain streams, Journal of Environmental Management, 295, 113066, ISSN: 0301-4797. DOI: 10.1016/j.jenvman.2021.113066.

Patel, C, Vadher, AN, Mathers, K, Dwyer, C, Wood, P (2021) Body size affects the vertical movement of benthic amphipods through subsurface sediments in response to drying, Hydrobiologia, 848, pp.1015-1025, ISSN: 0018-8158. DOI: 10.1007/s10750-020-04500-x.

Pledger, A, Brewin, P, Mathers, K, Phillips, J, Wood, P, Yu, D (2020) The effects of water injection dredging on low-salinity estuarine ecosystems: implications for fish and macroinvertebrate communities, Ecological Indicators, 122, 107244, ISSN: 1470-160X. DOI: 10.1016/j.ecolind.2020.107244.

Gething, KJ, Ripley, MC, Mathers, K, Chadd, RP, Wood, P (2020) The influence of substrate type on macroinvertebrate assemblages within agricultural drainage ditches, Hydrobiologia, 847, ISSN: 0018-8158. DOI: 10.1007/s10750-020-04416-6.

Mathers, K, White, JC, Guareschi, S, Hill, MJ, Heino, J, Chadd, R (2020) Invasive crayfish alter the long‐term functional biodiversity of lotic macroinvertebrate communities, Functional Ecology, 34(11), pp.2350-2361, ISSN: 0269-8463. DOI: 10.1111/1365-2435.13644.

Mathers, K, Michalec, F, Holzner, M, Weber, C (2020) Beneath the surface: Application of transparent super absorbent polymer substrates to track faunal activity within the sediment layer, Freshwater Biology, 65(11), pp.1923-1935, ISSN: 0046-5070. DOI: 10.1111/fwb.13588.

Beatty, C, Mathers, K, Patel, C, Constable, D, Wood, P (2020) Substrate mediated predator–prey interactions between invasive crayfish and indigenous and non-native amphipods, Biological Invasions, 22(9), pp.2713-2724, ISSN: 1387-3547. DOI: 10.1007/s10530-020-02292-8.

Mathers, K, White, JC, Fornaroli, R, Chadd, R (2020) Flow regimes control the establishment of invasive crayfish and alter their effects on lotic macroinvertebrate communities, Journal of Applied Ecology, 57(5), pp.886-902, ISSN: 0021-8901. DOI: 10.1111/1365-2664.13584.

McKenzie, M, Mathers, K, Wood, P, England, J, Foster, I, Lawler, D, Wilkes, M (2019) Potential physical effects of suspended fine sediment on lotic macroinvertebrates, Hydrobiologia, 847(3), pp.697-711, ISSN: 0018-8158. DOI: 10.1007/s10750-019-04131-x.

Mathers, KL, Stubbington, R, Leeming, D, Westwood, C, England, J (2019) Structural and functional responses of macroinvertebrate assemblages to long‐term flow variability at perennial and nonperennial sites, Ecohydrology, 12(6), ISSN: 1936-0584. DOI: 10.1002/eco.2112.

White, JC, Armitage, PD, Bass, JAB, Chadd, RP, Hill, MJ, Mathers, KL, Little, S, Wood, P (2019) How freshwater biomonitoring tools vary sub‐seasonally reflects temporary river flow regimes, River Research and Applications, 35(8), pp.1325-1337, ISSN: 1535-1459. DOI: 10.1002/rra.3501.

Hill, MJ, Mathers, KL, Little, S, Worrall, T, Gunn, J, Wood, P (2019) Ecological effects of a supra-seasonal drought on macroinvertebrate communities differ between near-perennial and ephemeral river reaches, Aquatic Sciences, 81, 62, ISSN: 1015-1621. DOI: 10.1007/s00027-019-0659-7.

Seddon, E, Hill, M, Greenwood, M, Mainstone, C, Mathers, K, White, JC, Wood, P (2019) The use of palaeoecological and contemporary macroinvertebrate community data to characterize riverine reference conditions, River Research and Applications, 35(8), pp.1302-1313, ISSN: 1535-1459. DOI: 10.1002/rra.3490.

Wilkes, MA, Gittins, JR, Mathers, KL, Mason, R, Casas‐Mulet, R, Vanzo, D, Mckenzie, M, Murray‐Bligh, J, England, J, Gurnell, A, Jones, JI (2019) Physical and biological controls on fine sediment transport and storage in rivers, WIREs Water, 6(2), ISSN: 2049-1948. DOI: 10.1002/wat2.1331.

Mathers, K, Rice, S, Wood, P (2019) Predator, prey and substrate interactions: the role of faunal activity and substrate characteristics, Ecosphere, ISSN: 2150-8925. DOI: 10.1002/ecs2.2545.

Rice, S, Pledger, A, Toone, J, Mathers, KL (2018) Zoogeomorphological behaviours in fish and the potential impact of benthic feeding on bed material mobility in fluvial landscapes, Earth Surface Processes andLandforms, 44(1), pp.54-66, ISSN: 0197-9337. DOI: 10.1002/esp.4541.

Mathers, K, Hill, MJ, Wood, CD, Wood, P (2018) The role of fine sediment characteristics and body size on the vertical movement of a freshwater amphipod, Freshwater Biology, ISSN: 0046-5070. DOI: 10.1111/fwb.13202.

Mathers, KL, Rice, S, Wood, P (2018) Discharge and suspended sediment time series as controls on fine sediment ingress into gravel river beds, Catena, 173, pp.253-263, ISSN: 0341-8162. DOI: 10.1016/j.catena.2018.10.001.

Clinton, KE, Mathers, K, Constable, D, Gerrard, C, Wood, P (2018) Substrate preferences of coexisting invasive amphipods, Dikerogammarus villosus and Dikerogammarus haemobaphes, under field and laboratory conditions, Biological Invasions, pp.1-10, ISSN: 1387-3547. DOI: 10.1007/s10530-018-1695-2.

Mathers, K, Collins, AL, England, J, Brierley, B, Rice, S (2017) The fine sediment conundrum; quantifying, mitigating and managing the issues, River Research and Applications, 33(10), pp.1509-1514, ISSN: 1535-1459. DOI: 10.1002/rra.3228.

Turley, MD, Bilotta, GS, Gasparrini, A, Sera, F, Mathers, KL, Humpheryes, I, England, J (Accepted for publication) The effects of non-native signal crayfish (Pacifastacus leniusculus) on fine sediment and sediment-biomonitoring, Science of the Total Environment, 601-602, pp.186-193, ISSN: 0048-9697. DOI: 10.1016/j.scitotenv.2017.05.106.

Harper, SE, Foster, IDL, Lawler, DM, Mathers, KL, Mckenzie, M, Petts, GE (Accepted for publication) The complexities of measuring fine sediment accumulation within gravel-bed rivers, River Research and Applications, ISSN: 1535-1459. DOI: 10.1002/rra.3198.

Mathers, K, Rice, S, Wood, P (2017) Temporal variability in lotic macroinvertebrate communities associated with invasive signal crayfish (Pacifastacus leniusculus) activity levels and substrate character, Biological Invasions, 20(3), ISSN: 1387-3547. DOI: 10.1007/s10530-017-1557-3.

Mathers, K, Hill, MJ, Wood, P (2017) Benthic and hyporheic macroinvertebrate distribution within the heads and tails of riffles during baseflow conditions, Hydrobiologia, ISSN: 0324-0924. DOI: 10.1007/s10750-017-3092-8.

Mathers, K, Rice, S, Wood, P (2017) Temporal effects of enhanced fine sediment loading on macroinvertebrate community structure and functional traits, Science of the Total Environment, ISSN: 0048-9697. DOI: 10.1016/j.scitotenv.2017.04.096.

Rice, S, Johnson, MF, Mathers, K, Reeds, J, Extence, CA (2016) The importance of biotic entrainment for base flow fluvial sediment transport, Journal of Geophysical Research - Earth Surface, 121(5), pp.890-906, ISSN: 2169-9011. DOI: 10.1002/2015JF003726.

Mathers, K, Chadd, RP, Dunbar, MJ, Extence, CA, Reeds, J, Rice, S, Wood, P (2016) The long-term effects of invasive signal crayfish (Pacifastacus leniusculus) on instream macroinvertebrate communities, Science of the Total Environment, 556, pp.207-218, ISSN: 0048-9697. DOI: 10.1016/j.scitotenv.2016.01.215.

Mathers, K, Chadd, RP, Extence, CA, Rice, S, Wood, P (2016) The implications of an invasive species on the reliability of macroinverterbrate biomonitoring tools used in freshwater ecological assessments, Ecological Indicators, ISSN: 1872-7034. DOI: 10.1016/j.ecolind.2015.11.051.

Hill, MJ, Death, RG, Mathers, K, Ryves, D, White, JC, Wood, P (2016) Macroinvertebrate community composition and diversity in ephemeral and perennial ponds on unregulated floodplain meadows in the UK, Hydrobiologia, pp.1-14, ISSN: 0018-8158. DOI: 10.1007/s10750-016-2856-x.

Mathers, K and Wood, P (2016) Fine sediment deposition and interstitial flow effects on macroinvertebrate community composition within riffle heads and tails, Hydrobiologia, ISSN: 0018-8158. DOI: 10.1007/s10750-016-2748-0.

Hill, M, Wood, P, Mathers, K (2015) The aquatic macroinvertebrate biodiversity of urban ponds in a medium-sized European town (Loughborough, UK), Hydrobiologia, ISSN: 0324-0924. DOI: 10.1007/s10750-015-2328-8.

Mathers, KL, Millett, J, Robertson, AL, Stubbington, R, Wood, PJ (2014) Faunal response to benthic and hyporheic sedimentation varies with direction of vertical hydrological exchange, Freshwater Biology, 59(11), pp.2278-2289, ISSN: 0046-5070. DOI: 10.1111/fwb.12430.

Mathers, KL and Wood, PJ (2012) The response of Gammarus pulex to fine sedimentation and vertical hydraulic exchange: An experimental investigation, Cave and Karst Science, 39(3), pp.123-126, ISSN: 1356-191X.

Mathers, K, Robinson, CT, Hill, M, Kowarik, C, Heino, J, Deacon, C, Weber, C (Accepted for publication) How effective are ecological metrics in supporting conservation and management in degraded streams?, Biodiversity and Conservation, ISSN: 0960-3115. DOI: 10.1007/s10531-024-02933-7.

Conferences

Mathers, K, Robinson, CT, Weber, C, Wood, P (2021) The importance of substrate heterogeneity and vertical hydrological connectivity for ecological diversity. In Wyss, CR, Cesare, GD, Lane, SN, Marti, C, Nitsche, M, Pauli, M, Schweizer, S, Weitbrecht, V (ed) International Symposium on Bedload Management 2021, Interlaken, Switzerland, pp.11-15, DOI: 10.3929/ethz-b-000513098.

Chapters

Wood, PJ, Armitage, PD, Hill, MJ, Mathers, KL, Millett, J (2016) Faunal response to fine sediment deposition in urban rivers. In Unknown Parent Title, Wiley, pp.219-238, ISBN: 9781119994343. DOI: 10.1002/9781118643525.ch11.

Wood, P, Armitage, PD, Hill, MJ, Mathers, K, Millett, J (2016) Faunal response to fine sediment deposition in urban rivers. In River Science: Research and Management for the 21st Century, © John Wiley & Sons, Ltd, pp.223-238, ISBN: 9781119994343. DOI: 10.1002/9781118643525.ch11.

Internet Publications

Mathers, K, Robinson, CT, Weber, C (2022) Supplementary Information files for: Patchiness in flow refugia use by macroinvertebrates following an artificial flood pulse.

Reports

Rice, SP, Mathers, K, Smith, J (2015) Biological control of invasive signal crayfish by European eel and associated effects on sediment dynamics and flood risk management, pp.1-33, Environment Agency.

Datasets

Mathers, K, Armitage, P, Bickerton, M, Hill, M, Mckenzie, M, Pardo, I, Tickner, D, Wood, P (2024) Macroinvertebrate data from sand, silt, gravel habitats in five English rivers, DOI: 10.17028/rd.lboro.26397802.

Snåre, H, García-Girón, J, Alahuhta, J, Bini, LM, Boda, P, Bonada, N, Brasil, LS, Callisto, M, Castro, DMP, Chen, K, Csabai, Z, Datry, T, Domisch, S, Márquez, JRG, Floury, M, Friberg, N, Gill, BA, González-Trujillo, JD, Göthe, E, Haase, P, Hamada, N, Hill, MJ, Hjort, J, Juen, L, Jupke, JF, de Faria, APJ, Li, Z, Ligeiro, R, Linares, MS, Luiza-Andrade, A, Macedo, DR, Mathers, K, Mellado-Díaz, A, Milosevic, D, Moya, N, Poff, NL, Rolls, RJ, Roque, FO, Saito, VS, Sandin, L, Schäfer, RB, Scotti, A, Siqueira, T, Martins, RT, Valente-Neto, F, Wang, BWJ, Xie, Z, Heino, J (2024) Supplementary information files for The relationships between biotic uniqueness and abiotic uniqueness are context dependent across drainage basins worldwide, DOI: 10.17028/rd.lboro.25586865.

Mckenzie, M, Scott, J, Wood, P, Mathers, K (2024) Supplementary information files for "Phenotypic plasticity of a Baetid mayfly larvae (Baetis rhodani) at sites with high levels of deposited fine sediment", DOI: 10.17028/rd.lboro.28229489.

Mathers, K, Wood, P, Mckenzie, M, Scott, J (2024) Ionocyte gill count from Baetis rhodani, DOI: 10.17028/rd.lboro.25233532.

Mckenzie, M, Brooks, A, Callisto, M, Collins, AL, Durkota, JM, Death, RG, Jones, JI, Linares, MS, Matthaei, CD, Monk, WA, Murphy, JF, Wagenhoff, A, Wilkes, M, Wood, P, Mathers, K (2023) Supplementary information files for Freshwater invertebrate responses to fine sediment stress: a multi‐continent perspective, DOI: 10.17028/rd.lboro.24885717.

Mathers, K, Armitage, P, Hill, M, Mckenzie, M, Pardo, I, Wood, P (2023) Seasonal habitat macroinvertebrate data from Mill Stream, Dorset, UK, DOI: 10.17028/rd.lboro.24132561.

Davis, NG, Mathers, K, Hodson, R, Matthaei, CD (2023) Supplementary information files for Monthly sampling reveals seasonal fine sediment fluctuations and riverine invertebrate community responses, DOI: 10.17028/rd.lboro.24658425.

Mathers, K, Brooks, A, Callisto, M, Collins, A, Durkota, J, Death, R, Jones, JI, Linares, M, Matthaei, C, Monk, W, Murphy, J, Wagenhoff, A, Wilkes, M, Wood, P, Mckenzie, M (2023) Paired visual fine sediment (<2mm) data and macroinvertebrate samples from Australia, Brazil, New Zealand and UK, DOI: 10.17028/rd.lboro.24615570.

Mathers, K, Armitage, PD, Hill, M, Mckenzie, M, Pardo, I, Wood, P (2023) Supplementary information files for "Seasonal variability of lotic macroinvertebrate communities at the habitat scale demonstrates the value of discriminating fine sediment fractions in ecological assessments", DOI: 10.17028/rd.lboro.28333316.

Mathers, K, Clinton, K, Constable, D, Gerrard, C, Patel, C, Wood, P (2023) Supplementary information files for "Invasion dynamics of Ponto‐Caspian amphipods leads to changes in invertebrate community structure and function", DOI: 10.17028/rd.lboro.28227920.

Mathers, K, Clinton, K, Constable, A, Gerrard, C, Patel, C, Wood, P (2023) Macroinvertebrate data from Pitsford Reservoir over three years, DOI: 10.17028/rd.lboro.23019596.

Mathers, K, Rice, S, Chadd, R, Wood, P (2022) Supplementary information files for Influence of invasive crayfish on fine sediment transport, ingress and bed storage in lowland rivers, DOI: 10.17028/rd.lboro.20368023.

Mathers, K, Doretto, A, Fenoglio, S, Hill, M, Wood, P (2022) Supplementary information files for Temporal effects of fine sediment deposition on benthic macroinvertebrate community structure, function and biodiversity likely reflects landscape setting, DOI: 10.17028/rd.lboro.19780225.

Mathers, K, Robinson, C, Weber, C (2021) Riverine fine sediment content and hyporheic physicochemistry and macroinvertebrate data associated with an artificial flood in Switzerland, DOI: 10.17028/rd.lboro.16434759.

Gething, KJ, Ripley, MC, Mathers, K, Chadd, RP, Wood, P (2020) Supplementary Information files for The influence of substrate type on macroinvertebrate assemblages within agricultural drainage ditches, DOI: 10.17028/rd.lboro.15173334.

Beatty, C, Mathers, K, Patel, C, Constable, D, Wood, P (2020) Supplementary Information files for: Substrate mediated predator-prey interactions between invasive crayfish and indigenous and non-native amphipods, DOI: 10.17028/rd.lboro.12821093.

Mathers, K, White, JC, Guareschi, S, Hill, MJ, Heino, J, Chadd, R (2020) Supplementary Information files for Invasive crayfish alter the long‐term functional biodiversity of lotic macroinvertebrate communities, DOI: 10.17028/rd.lboro.12800645.